Root development in young corn
Examples of seminal root damage include imbibitional chilling injury (Nielsen, 2012), post-germination injury from lethal or sub-lethal cold temperatures (Nielsen, 2012), and “salt” injury from excessive rates of starter fertilizer placed too close to the kernel. Symptoms of such root damage include retarded root elongation, brown tissue discoloration, prolific root branching, and outright death of root tissue. If the radicle root is damaged severely during its emergence from the kernel, the entire radicle root may die. Once the radicle has elongated a half-inch or so, damage to the root tip will not necessarily kill the entire root, but rather axillary root meristems may initiate extensive root branching in response to damage to the apical meristem.
The Nodal Root System
Nodal roots develop sequentially from individual nodes above the mesocotyl, beginning with the lowermost node in the area of the young seedling known as the "crown". Once a seedling has reached the V1 stage of development, one can usually identify the first set of nodal roots beginning to elongate from the lowermost node. By the V2 stage of development, the first set of nodal roots are clearly visible and the second set of nodal roots may be starting to elongate from the second node of the seedling. Each set or "whorl" of nodal roots begins to elongate from their respective nodes at about the same timing that each leaf collar emerges from the true whorl of the seedling.
Regarding Seeding Depth & Rooting Depth: Some folks believe that planting corn deeper encourages deeper rooting and vice versa. This belief is mostly myth with a slight hint of truth mixed in. It certainly is true that the depth of the seminal root system is influenced by seeding depth. However, the nodal root system that develops from the crown of the plant is not influenced much at all by seeding depth. This is because the depth of the crown is fairly constant regardless of seeding depth. During emergence of the seedling, the mesocotyl elongates and elevates the coleoptile and crown towards the soil surface. As the coleoptile nears the soil surface, changes in the ratio of red to far red wavelengths of light causes a change in the supply of one or more growth hormones from the coleoptile to the mesocotyl tissue and mesocotyl elongation consequently comes to a halt (Vanderhoef & Briggs, 1978). Since the depth at which the emerging seedling senses the change in red to far red light is fairly constant, the resulting depth of the crown (base) of the coleoptile is nearly the same (1/2 to 3/4 inch) for seeding depths of one inch or greater.
Elongation of the stalk tissue begins between leaf stages V4 and V5. Elongation of the internode above the fifth node usually elevates the sixth node above ground. Subsequent elongation of higher-numbered stalk internodes will result in higher and higher placement of the remaining stalk nodes. Sets of nodal roots that form at above ground stalk nodes are commonly referred to as “brace” roots, but function identically to those nodal roots that form below ground. If surface soil conditions are favorable (moist and not excessively hot), brace roots will successfully penetrate the soil, proliferate, and effectively scavenge the upper soil layers for water and nutrients.
Corny Trivia:Root hairs are lateral extensions of root epidermal cells, grow to a length of several millimeters, and number about 200 per sq. millimeter (Gardner et al., 1985). Their typical life span is only about 2 days at moderate temperatures and less so at higher temperatures (Gardner et al., 1985). Root hairs are visible even on the radicle root of a young seedling. Collectively, the surface area represented by root hairs is very large and can account for a large share of nutrient and moisture uptake by the plant.
Corny Trivia: The primary meristem of a root is located near the root tip. Elongation of cells behind the meristem leads to elongation of the root.
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